The lons variable doesn't seem to be defined in the step1_preprocess_forcings file. I'm not exactly sure what it's supposed to be either. (Should it also be a vector like lats is currently defined? Or something else?)
Full error message follows.
Operation terminated by user during step1_preprocess_forcings (line 83)
Error using save
Variable 'lons' not found.
Error in step1_preprocess_forcings (line 109)
save processed_forcings/cesm_lons.mat lons